the article has many different typefaces with a sans serif possibly being used most as its used for the body text.
then second most used has to be the script type font they used for the main title.
and coming in last would have to be the tradisional (lot of thick and thin contrast) serif font(s) they uses as subheadings and for extra lil notes as well as the drive name (mss folio)
i also noticed they used italics all the way at the bottom.
i think the legibility of the article is only ok with all the different fonts and hidden text but that’s also why i think it is so readable because it’s fun like your going on a little type scavenger hunt
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